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    Analogies between the virial theorem and the Price equation

    Catherine Felce1, Steinunn Liorsdóttir2, and Lior Pachter3,*

    • 1Division of Physics, Mathematics and Astronomy, California Institute of Technology, Pasadena, California 91125, USA
    • 2Flintridge Preparatory School, La Cañada, California 91011, USA
    • 3Division of Biology and Biological Engineering, and Department of Computing and Mathematical Sciences, California Institute of Technology, Pasadena, California 91125, USA

    • *Contact author: lpachter@caltech.edu

    Phys. Rev. E 112, 054139 – Published 21 November, 2025

    DOI: https://doi.org/10.1103/r8bd-lhm3

    Abstract

    We observe that the time-averaged continuous Price equation is identical to the positive momentum virial theorem, and we discuss the applications and implications of this connection. We also introduce ecological models using a maternal effect, which can describe arbitrary population size cycles and trait evolution across spatially separated populations. The virial theorem sheds light on the time-averaged behavior of these models.

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