Trevor K. GrandPre^20,21, Alexander Y. Grosberg²², Ivar S. Haugerud²³, William M. Jacobs²⁴, Jerelle A. Joseph^1,2,*, Frank Jülicher^25,†, Kurt Kremer²⁶, Guido Kusters²⁷, Liedewij Laan²⁸, Keren Lasker²⁹, Kathrin S. Laxhuber²⁵, Hyun O. Lee^16,‡, Kathy F. Liu^{30,31,32,33,34}, Dimple Notani³⁵, Yicheng Qiang²⁷, Paul Robustelli³⁶, Leonor Saiz³⁷, Omar A. Saleh³⁸, Helmut Schiessel³⁹, Jeremy D. Schmit⁴⁰, Meng Shen⁴¹, Krishna Shrinivas^42,43, Antonia Statt⁴⁴, Andres R. Tejedor^17,18,19, Tatjana Trcek⁴⁵, Christoph A. Weber²³, Stephanie C. Weber^46,47,§, Ned S. Wingreen^48,49,∥, Huaiying Zhang⁵⁰, Yaojun Zhang^51,52, Huan-Xiang Zhou⁵³, and David Zwicker^27,¶

• ¹Department of Chemical and Biological Engineering, Princeton University, New Jersey 08544, USA
• ²Omenn-Darling Bioengineering Institute, Princeton University, New Jersey 08544, USA
• ³Institute of Numerical Mathematics and Optimization, TU Bergakademie Freiberg, 09599 Freiberg, Germany
• ⁴Faculty of Informatics/Mathematics, HTW Dresden, 01069 Dresden, Germany
• ⁵Department of Physics, King's College London, London WC2R 2LS, United Kingdom
• ⁶Department of Molecular and Human Genetics, Baylor College of Medicine, Houston, Texas 77030, USA
• ⁷Jan and Dan Duncan Neurological Research Institute, Texas Children's Hospital, Houston, Texas 77030, USA
• ⁸Therapeutic Innovation Center (THINC), Baylor College of Medicine, Houston, Texas 77030, USA
• ⁹Center for Alzheimer's and Neurodegenerative Diseases (CAND), Baylor College of Medicine, Houston, Texas 77030, USA
• ¹⁰Dan L Duncan Comprehensive Cancer Center (DLDCCC), Baylor College of Medicine, Houston, Texas 77030, USA
• ¹¹Department of Biochemistry and Molecular Biology, Johns Hopkins Bloomberg School of Public Health, Baltimore, Maryland 21205, USA
• ¹²Department of Biophysics and Biophysical Chemistry, Johns Hopkins School of Medicine, Baltimore, Maryland 21205, USA
• ¹³Department of Oncology, Johns Hopkins School of Medicine, Baltimore, Maryland 21205, USA
• ¹⁴Department of Biomedical, Metabolic and Neural Sciences, University of Modena and Reggio Emilia, Modena 41125, Italy
• ¹⁵Department of Biology, Massachusetts Institute of Technology, Cambridge, Massachusetts 02139, USA
• ¹⁶Department of Biochemistry, University of Toronto, Toronto, Ontario M5S 1A8, Canada
• ¹⁷Department of Physical Chemistry, University Complutense of Madrid, Madrid 28040, Spain
• ¹⁸Instituto Pluridisciplinar, Universidad Complutense de Madrid, P.º de Juan XXIII, 1, Moncloa - Aravaca, 28040 Madrid, Spain
• ¹⁹Yusuf Hamied Department of Chemistry, University of Cambridge, Lensfield Road, Cambridge CB2 1EW, United Kingdom
• ²⁰Department of Physics and Center for Biomolecular Condensates, Washington University in St. Louis, St. Louis, Missouri 63130, USA
• ²¹National Institute for Theory and Mathematics in Biology, Northwestern University and The University of Chicago, Chicago, Illinois 60611, USA
• ²²Department of Physics and Center for Soft Matter Research, New York University, 726 Broadway, New York, New York 10003, USA
• ²³Faculty of Mathematics, Natural Sciences, and Engineering, Institute of Physics, University of Augsburg, Universitätsstrasse 1, 86159 Augsburg, Germany
• ²⁴Department of Chemistry, Princeton University, Princeton, New Jersey 08544, USA
• ²⁵Max Planck Institute for the Physics of Complex Systems, Nöthnitzer Str. 38, 01187 Dresden, Germany
• ²⁶Max Planck Institute for Polymer Research, Ackermannweg 10, 55128 Mainz, Germany
• ²⁷Max Planck Institute for Dynamics and Self-Organization, Am Faßberg 17, 37077 Göttingen, Germany
• ²⁸Department of Bionanoscience, Kavli Institute of Nanoscience Delft, Delft University of Technology, Delft, The Netherlands
• ²⁹Department of Integrative Structural and Computational Biology, The Scripps Research Institute, La Jolla, California 92037, USA
• ³⁰Department of Biochemistry and Biophysics, Perelman School of Medicine, University of Pennsylvania, Philadelphia, Pennsylvania 19104, USA
• ³¹Graduate Group in Biochemistry and Molecular Biophysics, Perelman School of Medicine, University of Pennsylvania, Philadelphia, Pennsylvania 19104, USA
• ³²Abramson Family Cancer Research Institute, Perelman School of Medicine, University of Pennsylvania, Philadelphia, Pennsylvania 19104, USA
• ³³Penn Institute for RNA Innovation, University of Pennsylvania, Philadelphia, Pennsylvania 19104, USA
• ³⁴Penn Center for Genome Integrity, University of Pennsylvania, Philadelphia, Pennsylvania 19104, USA
• ³⁵National Centre for Biological Sciences-TIFR, GKVK Campus, Bellary Road, Bangalore 560065, India
• ³⁶Department of Chemistry, Dartmouth College, Hanover, New Hampshire 03755, USA
• ³⁷Department of Biomedical Engineering, University of California, Davis, California 95616, USA
• ³⁸Materials and Physics Departments, University of California, Santa Barbara, California 93110, USA
• ³⁹Cluster of Excellence Physics of Life, TU Dresden, 01062 Dresden, Germany
• ⁴⁰Department of Physics, Kansas State University, Manhattan, Kansas 66506, USA
• ⁴¹Department of Physics, California State University, Fullerton, California 92831, USA
• ⁴²Department of Chemical and Biological Engineering, Northwestern University, Evanston, Illinois 60208, USA
• ⁴³Center for Synthetic Biology, Northwestern University, Evanston, Illinois 60208, USA
• ⁴⁴Department of Materials Science and Engineering, University of Illinois Urbana-Champaign, Urbana, Illinois 61801, USA
• ⁴⁵Department of Biology, Johns Hopkins University, Baltimore, Maryland 21218, USA
• ⁴⁶Department of Biology, McGill University, Montreal, H3A 1B1, Canada
• ⁴⁷Department of Physics, McGill University, Montreal H3A 2T8, Canada
• ⁴⁸Department of Molecular Biology, Princeton University, New Jersey 08544, USA
• ⁴⁹Lewis-Sigler Institute for Integrative Genomics, Princeton University, New Jersey 08544, USA
• ⁵⁰Department of Biological Sciences, Carnegie Mellon University, Pittsburgh, Pennsylvania, 15213, USA
• ⁵¹Department of Physics & Astronomy, Johns Hopkins University, Baltimore, Maryland 21218, USA
• ⁵²Department of Biophysics, Johns Hopkins University, Baltimore, Maryland 21218, USA
• ⁵³Department of Chemistry and Department of Physics, University of Illinois Chicago, Chicago, Illinois 60607, USA

• ^*Contact author: jerellejoseph@princeton.edu
• ^†Contact author: julicher@pks.mpg.de
• ^‡Contact author: hyunokate.lee@utoronto.ca
• ^§Contact author: steph.weber@mcgill.ca
• ^∥Contact author: wingreen@princeton.edu
• ^¶Contact author: david.zwicker@ds.mpg.de